Author Contributions
Y.-S.J., L.S. developed the idea of this work. Y.-S.J., L.S. designed the experiments. L.S. performed the experiments. L.S., C.A., Y.-G.L. and Y.-S.J. wrote the manuscript.
Funding This work was funded by the DOE Center for Advanced Bioenergy and Bioproducts Innovation (U.S. Department of Energy, Office of Science, Office of Biological and Environmental Research under Award Number DE-SC0018420). Any opinions, findings, and conclusions or recommendations expressed in this publication are those of the author(s) and do not necessarily reflect the views of the U.S. Department of Energy. L. S. would like to thank the China Scholarship Council (CSC) for financial support.