Acknowledgements
This work was supported by the Fund of Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang) (ZJW-2019-06), Youth Talent Program Supported by Laboratory for Marine Fisheries Science and Food Production Processes, Pilot National Laboratory for Marine Science and Technology (Qingdao) (2018-MFS-T08), Shandong Superior Variety Project (2016LZGC009),the AoShan Talents Cultivation Program Supported by Pilot National Laboratory for Marine Science and Technology (Qingdao) [2017ASTCP-OS15] and Shandong Taishan Scholar Climbing Project.