Author contributions
S.L.C., H.R.L and C.H.Z. conceived the project. S.L.C. and C.H.Z. managed the project. Q.Z., H.X. and C.X.T. collected the sequencing samples. X.Y.G. extracted the genomic DNA and performed the genome sequencing. Q.Z., G.H and Y. H. analyzed the data. Q.Z. wrote the manuscript. All authors reviewed and approved the final manuscript.