Sequential Sum-of-Squares where the impact of each variable is considered after the impact of parasite species is accounted. Grey shading indicates significant impact without violating assumptions of PERMANOVA. The PERMANOVA p-value (top; *=p<0.05, **=p<0.005, ***=p<0.0005), R2 (middle) and p-value for homoscedasticity (bottom, significance indicates violation of the assumptions of PERMANOVA) are provided for each dataset.