Acknowledgements

First, we would like to thank Michele Esposito, from CASC, for her enduring support with databasing, imaging processing, proofreading, and her overall support in the lab. The fieldwork that provided the specimens used in this study could not have been completed without the help of the Ministry of Environment and Forest, the Madagascar National Parks, and the Parc Botanique et Zoologique de Tsimbazaza, and the gracious support of the Malagasy people. The authors wish to thank Dr. Bethan Davies for her permission to use the graphics in Figure 1.
This study was supported by the National Science Foundation under Grant No. DEB-0072713, DEB-0344731, and DEB-0842395. Finally, SC was partly supported by the Schlinger Fellowship at the California Academy of Sciences.
The ant specimens used in this study comply with the regulations for export and exchange of research samples outlined in the Convention on Biological Diversity and the Convention on International Trade in Endangered Species of Wild Fauna and Flora. For field work conducted in Madagascar, permits to research, collect, and export ants were obtained from the Ministry of Environment and Forest as part of an ongoing collaboration between the California Academy of Sciences and the Ministry of Environment and Forest, Madagascar National Parks and Parc Botanique et Zoologique de Tsimbazaza (Approval Numbers: N° 0142N/EA03/MG02, N° 340N-EV10/MG04, N° 69 du 07/04/06, N° 065N-EA05/MG11, N° 047N-EA05/MG11, N° 083N-A03/MG05, N° 206 MINENVEF/SG/DGEF/DPB/SCBLF, N° 0324N/EA12/MG03, N° 100 l\fEF/SG/DGEF/DADF/SCBF, N° 0379N/EA11/MG02, N° 200N/EA05/MG02). Authorization for export was provided by the Director of Natural Resources.