Acknowledgements
FY was supported by the National Science Foundation of China (NSFC) grant U1904105; HZ by the National Science Foundation of China (NSFC) grant 3217030369; and XY by the National Science Foundation of China (NSFC) grant 32070447 and the Young Talents Invitation Program of Shandong Provincial Colleges and Universities grant 20190601. Thanks are given to Bo Wang for data analysis of phylogenetic partial R2s.