Ethical Statement
Capture, handling, and voucher collection complied with the guidelines recommended by the American Society of Mammalogists (Gannon and Sikes, 2007), and small mammal sampling guidelines compiled by Aegerter and Heritage (2005) and Kunz and Parsons (2009); and were approved by the Science Faculty Animal Ethics Committee at the University of Cape Town (Clearance Number 2013/2011/V6/DJ). All workers handling bats were vaccinated for rabies and were required to use protective gloves when handling bats and samples.