Authors’ Contributions

MD conceived the presented ideas in discussion with TM. MD designed the model and computational framework with assistance from LS and TU. MD and MG wrote the manuscript with support from MW, ES, and TM. The camera trap project in the present study was made possible by BH, CS, LA, MG, and TM; with field assistance and data collection provided by all authors. TM secured funding provided by BH, CS, and LA. TM and MG supervised the project. All authors contributed to the drafts of the publication.