Landscape of the study area
The state space of the study area was defined by buffering the minimum area rectangle of the sample units by 1000 meters, producing a state-space of 10.4 km2. Agricultural areas comprised 76% (548 of 725 pixels), coniferous forests 12% (88 pixels), mixed forests 6% (45 pixels) and transitional woodland 6% (44 pixels) (Figure 1). 51.1% of the plots (47 of 92 plots) were coniferous forest, 30.4% (28 plots) mixed forest and 18.5% (17 plots) transitional woodland.