AUTHOR CONTRIBUTIONS
D.W. X.Z., C.C. whole-genome sequencing data analysis and discussed the results and prepared the manuscript. X.Z. X.Y. and S.C. drought stress experiments and transcriptome data analysis. L.S., X.F., H.K., Y.W., and Y.C. provide experimental guidance. Y.W., Y.Z., and D.L. provide data analysis guidance. J.L., D.L., W.Z., and Y.Z. discussed the manuscript. D.W., H.Z. review and editing the manuscript. H.Z. funding acquisition. Y.Z. and H.Z. designed and supervised this study.