Acknowledgements
We would like to express our sincere gratitude to Ms. Miwako Sugawara and Mr. Shinjiro Itoh, and all members in the laboratory of Wildlife Biology and Medicine, Hokkaido University for their efforts of DNA analysis. We wish to thank Hatsusaburo Ose and all the members of the Shiretoko Fishery Productive Association for their generous support. We thank all the members of the Shiretoko Nature Foundation and South Shiretoko Brown Bear Information Center for their generous support. We also thank all the people involved in sample collection. This work was supported by AIR DO Co., Ltd, Daikin Industries, Ltd., Mitsui & Co. Environment Fund (R14-0026), Fuji Film Green Fund, a Grant-in-Aid for Scientific Research from Japan Society for the Promotion of Science (19K06833), and the Environment Research and Technology Development Fund (4-1905; JPMEERF20194005) of the Environmental Restoration and Conservation Agency of Japan. This work was supported by the World-leading Innovative and Smart Education (WISE) Program (1801) from the Ministry of Education, Culture, Sports, Science, and Technology, Japan.