GPS tracking data analysis
To obtain the distances traveled per hour (speed), we divided the distance traveled between two consecutive positions by the time lapse, with a maximum of two hours. We used the Kruskal-Wallis test (data failed a test for normality) to examine if there were differences relative to time of day and the Games-Howell test (Zar, 1999) to identify period of peak activity. We also used the nonparametric Mann-Whitney U-test to evaluate travel differences between sexes and the dry (December - May) and rainy (June- November) seasons. All statistical analyses were performed with IBM SPSS Statistics ver. 22.0. Significance level was set at P < 0.05 and we present data as mean ± standard deviation (SD).