References
Adams, B. J., Schnitzer, S. A., and Yanoviak, S. P. 2017. Trees as islands: canopy ant species richness increases with the size of liana-free trees in a Neotropical forest. Ecography 40 : 1067–1075.
Adams, B. J., Schnitzer, S. A., and Yanoviak, S. P. 2019. Connectivity explains local ant community structure in a Neotropical forest canopy: a large‐scale experimental approach. Ecology 100 : e02673.
Antoniazzi, R., Viana-Junior, A. B., Pelayo-Martínez, J., Ortiz-Lozada, L., Neves, F. S., Leponce, M., and Dáttilo, W. 2021. Distance–decay patterns differ between canopy and ground ant assemblages in a tropical rainforest. J. Trop. Ecol. 1-9.
Ashton, L. A., Nakamura, A., Basset, Y., Burwell, C. J., Cao, M., Eastwood, R., … Kitching, R. L. 2016. Vertical stratification of moths across elevation and latitude. Journal of Biogeogr.43: 59–69.
Baselga, A., and Orme, C. D. L. 2012. betapart: an R package for the study of beta diversity. Methods ecol evol. 3 : 808-812.
Basham, E. W., Seidl, C. M., Andriamahohatra, L. R., Oliveira, B. F., and Scheffers, B. R. 2018. Distance–decay differs among vertical strata in a tropical rainforest. J. Anim. Ecol 88 : 114-124
Basset, Y., Cizek, L., Cuénoud, P., Didham, R. K., Guilhaumon, F., Missa, O., … Leponce, M. 2012. Arthropod diversity in a tropical forest. Science . 338: 1481-1484.
Bélisle, M. 2005. Measuring landscape connectivity: The challenge of behavioral landscape ecology. Ecology 88: 1988–1995.
Blanchet, F. G., Legendre, P., and Borcard, D. 2008. Forward selection of explanatory variables. Ecology  89 : 2623-2632.
Borowiec, M. L. 2016. Generic revision of the ant subfamily dorylinae (hymenoptera, formicidae). ZooKeys 608 : 1
Brown, N. 1993. The implications of climate and gap microclimate for seedling growth conditions in a Bornean lowland rain forest. J Trop. Ecol. 153-168.
Bütikofer, L., Anderson, K., Bebber, D. P., Bennie, J. J., Early, R. I., and Maclean, I. M. D. 2020. The problem of scale in predicting biological responses to climate. Glob. Chang. Biol.26 : 6657-6666.
Chesters, D., Beckschäfer, P., Orr, M. C., Adamowicz, S. J., Chun, K. P., and Zhu, C. D. 2019. Climatic and vegetational drivers of insect beta diversity at the continental scale. Ecol. Evol. 9 : 13764-13775.
Cottenie, K. 2005. Integrating environmental and spatial processes in ecological community dynamics. Ecol Lett. 8 : 1175–1182.
Daily, G. C., and Ehrlich, P. R. 1995. Preservation of biodiversity in small rainforest patches: rapid evaluations using butterfly trapping. Biodivers. Conserv . 4 : 35-55
Dáttilo, W., Fagundes, R., Gurka, C. A., Silva, M. S., Vieira, M. C., Izzo, T. J., … and Rico-Gray, V. 2014. Individual-based ant-plant networks: diurnal-nocturnal structure and species-area relationship. PLoS One.  9 : e99838.
Davies, A. B., and Asner, G. P. 2014. Advances in animal ecology from 3D-LiDAR ecosystem mapping. Trends Ecol. Evol.29 : 681–691.
Davis, K. T., Dobrowski, S. Z., Holden, Z. A., Higuera, P. E., and Abatzoglou, J. T. 2019. Microclimatic buffering in forests of the future: the role of local water balance. Ecography , 42 : 1–11.
Davidson, D. W. 1997. The role of resource imbalances in the evolutionary ecology of tropical arboreal ants. Biol. J. Linn. Soc. 61 : 153-181.
De Frenne, P., Zellweger, F., Rodríguez-Sánchez, F., Scheffers, B. R., Hylander, K., Luoto, M., … Lenoir, J. 2019. Global buffering of temperatures under forest canopies. Nature Ecol Evol. 3 : 744–749.
Dial, R., Bloodworth, B., Lee, A., Boyne, P. and Heys, J., 2004a. The distribution of free space and its relation to canopy composition at six forest sites. Forest Science 50 : 312-325.
Dial, R. J., Sillett, S. C., Antoine, M. E., and Spickler, J. C. 2004b. Methods for horizontal movement through forest canopies. Selbyana25:151-163.
Dial, R. J., Ellwood, M. D. F., Turner, E. C., and Foster, W. A. 2006. Arthropod Abundance, Canopy Structure, and Microclimate in a Bornean Lowland Tropical Rain Forest. Biotropica. 38 : 643–652.
Dial, R. J., Nadkarni, N. M., and Jewell III, C. D. 2011. Canopy structure in a 650-year Douglas-fir chronosequence in western Washington: Distribution of canopy elements and open space. Forest Science 57 : 309-319.
Dorow, W.H., and Kohout, RJ. 1995. A review of the subgenus Hemioptica Roger of the genus Polyrhachis Fr. Smith with description of a new species (Hymenoptera: Formicidae: Formicinae). Zoologische Mededelingen . 69 : 93-104.
Ehrlich, P. R., and Wilson, E. O. 1991. Biodiversity studies: Science and policy. Science , 253 : 758–762.
Fayle, T. M., Turner, E. C., Snaddon, J. L., Chey, V. K., Chung, A. Y. C., Eggleton, P., and Foster, W. A. 2010. Oil palm expansion into rain forest greatly reduces ant biodiversity in canopy, epiphytes and leaf-litter. Basic Appl Ecol. 11 : 337-345.
Fayle, T. M., Yusah, K. M., and Hashimoto, Y. 2014. Key to the ant genera of Borneo in English and Malay. http://www. tomfayle. com/Ant% 20key. htm.
Fetcher, N., Oberbauer, S. F., and Strain, B. R. 1985. Vegetation effects on microclimate in lowland tropical forest in Costa Rica.Int. J. Biometeorol . 29 : 145–155.
Floren, A., Wetzel, W., and Staab, M. 2014. The contribution of canopy species to overall ant diversity (Hymenoptera: Formicidae) in temperate and tropical ecosystems. Myrmecol. News 19 : 65-74.
Goslee, S. C., and Urban, D. L. 2007. The ecodist package for dissimilarity-based analysis of ecological data. J. Stat. Softw . 22: 1-19.
Hardwick, S. R., Toumi, R., Pfeifer, M., Turner, E. C., Nilus, R., and Ewers, R. M. 2015. The relationship between leaf area index and microclimate in tropical forest and oil palm plantation: Forest disturbance drives changes in microclimate. Agric For Meteorol201 : 187-195.
Huang, G., and Catterall, C. P. 2021. Effects of habitat transitions on rainforest bird communities across an anthropogenic landscape mosaic.Biotropica . 53 : 130-141.
Hung AC. 1970 A revision of ants of the subgenus Polyrhachis Fr. Smith (Hymenoptera: Formicidae: Formicinae). Orient. Insects.4 :1-36.
Jarron, L. R., Coops, N. C., MacKenzie, W. H., Tompalski, P., & Dykstra, P. 2020. Detection of sub-canopy forest structure using airborne LiDAR. Remote Sens. Environ. 244 : 111770.
Jorda, G., Marbà, N., Bennett, S., Santana-Garcon, J., Agusti, S., and Duarte, C. M. 2020. Ocean warming compresses the three-dimensional habitat of marine life. Nature Ecol. Evol. 4 : 109–114.
Kaspari, M. 1993. Body size and microclimate use in Neotropical granivorous ants. Oecologia . 96 : 500-507.
Kaspari, M., Clay, N. A., Lucas, J., Yanoviak, S. P., and Kay, A. 2015. Thermal adaptation generates a diversity of thermal limits in a rainforest ant community. Global Change Biol. 21 : 1092–1102.
Kaspari, M., and Yanoviak, S. P. 2001. Bait Use in Tropical Litter and Canopy Ants—Evidence of Differences in Nutrient Limitation1.Biotropica . 33 : 207-211.
Klimes, P. 2017. Diversity and specificity of ant-plant interactions in canopy communities: insights from primary and secondary tropical forests in New Guinea. Ant-Plant Interactions–Impacts of Humans on Terrestrial Ecosystems , 26-51.
Kitching, R. L., Ashton, L. A., Nakamura, A., Whitaker, T., and Khen, C. V. 2013. Distance-driven species turnover in Bornean rainforests: Homogeneity and heterogeneity in primary and post-logging forests.Ecography . 36 : 675-682.
Kohout, R.J. 2006a. Review of Polyrhachis (Cyrtomyrma) Forel (Hymenoptera: Formicidae: Formicinae) of Australia, Borneo, New Guinea and the Solomon Islands with descriptions of new species.Memoirs-Queensland Museum , 52: 87.
Kohout R.J. 2006b. A review of the Polyrhachiscryptoceroides species-group with description of a new species from Thailand (Hymenoptera: Formicidae). Myrmecologische Nachrichten .;8:145-50.
Klimes, P., Idigel, C., Rimandai, M., Fayle, T. M., Janda, M., Weiblen, G. D., and Novotny, V. 2012. Why are there more arboreal ant species in primary than in secondary tropical forests? J. Anim. Ecol.81 : 1103–1112.
Law, S. J., Bishop, T. R., Eggleton, P., Griffiths, H., Ashton, L., and Parr, C. 2020. Darker ants dominate the canopy: Testing macroecological hypotheses for patterns in colour along a microclimatic gradient.J. Anim. Ecol. 89: 347–359.
Lessard, J. P., Dunn, R. R., and Sanders, N. J. 2009. Temperature-mediated coexistence in temperate forest ant communities.Insectes Soc. 56 : 149-156.
Li, G., Cheng, L., Zhu, J., Trenberth, K. E., Mann, M. E., & Abraham, J. P. (2020). Increasing ocean stratification over the past half-century. Nature Climate Change 10 : 1116-1123.
MacArthur, R. H., and Horn, H. S. 1969. Foliage profile by vertical measurements. Ecology . 50 : 802-804.
Majer, J. D. 1993. Comparison of the arboreal ant mosaic in Ghana, Brazil, Papua New Guinea and Australia-its structure and influence on arthropod diversity. Hymenoptera and Biodiversity. 115-141.
Malhi, Y., Doughty, C., and Galbraith, D. 2011. The allocation of ecosystem net primary productivity in tropical forests. Philos. Trans. R. Soc. B: Biol. Sci.366 : 3225-3245.
Matthiessen, B., Mielke, E., and Sommer, U. 2010. Dispersal decreases diversity in heterogeneous metacommunities by enhancing regional competition. Ecology. 91 : 2022-2033.
Nakamura, A., Kitching, R. L., Cao, M., Creedy, T. J., Fayle, T. M., Freiberg, M., … Ashton, L. A. 2017. Forests and Their Canopies: Achievements and Horizons in Canopy Science. Trends Ecol. Evol. 32 : 438–451.
Nekola, J. C., and McGill, B. J. 2014. Scale dependency in the functional form of the distance decay relationship. Ecography .37 : 309–320.
Novotny, V., Miller, S. E., Hulcr, J., Drew, R. A. I., Basset, Y., Janda, M., … Weiblen, G. D. 2007. Low beta diversity of herbivorous insects in tropical forests. Nature 448 : 692–695.
Oliveira, B. F., and Scheffers, B. R. 2019) Vertical stratification influences global patterns of biodiversity. Ecography .42 : 249-249.
Oksanen, J., Blanchet, F. G., Kindt, R., Legendre, P., Minchin, P. R., O’hara, R. B., … and Wagner, H. 2013. Community ecology package.R package version , 2-0.
Parker, G. G. 1995. Structure and microclimate of forest canopies. In M.D. Lowman, and N. M. Nadkarni (Eds.), Forest canopies (pp. 73–106). SanDiego, CA: Academic Press Inc.
Parr, C. L., and Gibb, H. 2010. Competition and the role of dominant ants. Ant ecology , 77-96.
Perfecto, I., and Vandermeer, J. 1996. Microclimatic changes and the indirect loss of ant diversity in a tropical agroecosystem. Oecologia108 : 577-582.
Plowman, N. S., Mottl, O., Novotny, V., Idigel, C., Philip, F. J., Rimandai, M., and Klimes, P. 2019. Nest microhabitats and tree size mediate shifts in ant community structure across elevation in tropical rainforest canopies. Ecography . 43: 431-442.
Powell, S., Costa, A. N., Lopes, C. T., and Vasconcelos, H. L. 2011. Canopy connectivity and the availability of diverse nesting resources affect species coexistence in arboreal ants. J. Anim. Ecol.80 : 352-360
R core team. 2013. R: A language and environment for statistical computing.
Ramette, A., and Tiedje, J. M. 2007. Multiscale responses of microbial life to spatial distance and environmental heterogeneity in a patchy ecosystem. Proc. Natl. Acad. Sci. U.S.A. 104 :2761–2766.
Reaka-Kudla, M.L. 1997. The Global Biodiversity of Coral Reefs: A Comparison with Rain Forests. Biodiversity II: Understanding and Protecting Our Biological Resources . Joseph Henry Press.
Ribas, C. R., and Schoereder, J. H. 2007. Ant communities, environmental characteristics and their implications for conservation in the Brazilian Pantanal. Biodivers. Conserv . 16 : 1511-1520
Roisin, Y., Dejean, A., Corbara, B., Orivel, J., Samaniego, M., and Leponce, M. 2006. Vertical stratification of the termite assemblage in a neotropical rainforest. Oecologia , 149 : 301–311.
Sanders, N. J., Crutsinger, G. M., Dunn, R. R., Majer, J. D., and Delabie, J. H. 2007. An ant mosaic revisited: Dominant ant species disassemble arboreal ant communities but co‐occur randomly. Biotropica39 : 422-427.
Scheffers, B. R., Edwards, D. P., Macdonald, S. L., Senior, R. A., Andriamahohatra, L. R., Roslan, N., … Williams, S. E. 2017. Extreme thermal heterogeneity in structurally complex tropical rain forests. Biotropica 49 : 35–44.
Scheffers, B. R., Phillips, B. L., Laurance, W. F., Sodhi, N. S., Diesmos, A., and Williams, S. E. 2013. Increasing arboreality with altitude: A novel biogeographic dimension.Proc. Royal Soc. B ,1770: 20131581.
Schödl, S. 1998. Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae). Annalen des Naturhistorischen Museums in Wien. Serie B für Botanik und Zoologie,361-394.
Sheehan, T. N., Ulyshen, M. D., Horn, S., and Hoebeke, E. R. 2019. Vertical and horizontal distribution of bark and woodboring beetles by feeding guild: is there an optimal trap location for detection? J. Pest. Science. 92 : 327–341.
Shenkin, A., Chandler, C. J., Boyd, D. S., Jackson, T., Disney, M., Majalap, N., … Malhi, Y. 2019. The World’s Tallest Tropical Tree in Three Dimensions. Front. for. glob. change. 2 : 32.
Soininen, J., Heino, J., and Wang, J. 2018. A meta-analysis of nestedness and turnover components of beta diversity across organisms and ecosystems. Global Ecol. Biogeogr. 27 : 96–109.
Southwood, T. R. E., and Kennedy, C. E. J. 1983. Trees as Islands.Oikos . 359-371
Sreekar, R., Corlett, R. T., Dayananda, S., Goodale, U. M., Kilpatrick, A., Kotagama, S. W., … and Goodale, E. 2017. Horizontal and vertical species turnover in tropical birds in habitats with differing land use.Biol. Lett. 13 : 20170186.
Sreekar, R., Koh, L. P., Mammides, C., Corlett, R. T., Dayananda, S., Goodale, U. M., … Goodale, E. 2020. Drivers of bird beta diversity in the Western Ghats–Sri Lanka biodiversity hotspot are scale dependent: roles of land use, climate, and distance. Oecologia .193 : 801-809.
Stark, A. Y., Adams, B. J., Fredley, J. L., & Yanoviak, S. P. (2017). Out on a limb: Thermal microenvironments in the tropical forest canopy and their relevance to ants. J. Therm. Biol. 69 , 32-38.
Venegas‐Li, R., Levin, N., Possingham, H., and Kark, S. 2018. 3D spatial conservation prioritisation: Accounting for depth in marine environments. Methods. Ecol. Evol. 9 : 773–784.
Walsh, R. P. D., and Newbery, D. M. 1999. The ecoclimatology of Danum, Sabah, in the context of the world’s rainforest regions, with particular reference to dry periods and their impact. Proc. Royal Soc. B.354 : 1869-1883
Ward, P. S., Blaimer, B. B., and Fisher, B. L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the generaColobopsis and Dinomyrmex . Zootaxa . 4072 : 343-357
Ward, P. S., Brady, S. G., Fisher, B. L., and Schultz, T. R. 2015. The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Syst. Entomol.40 : 61-81.
Weiss, M., Procházka, J., Schlaghamerský, J., and Cizek, L. 2016. Fine-Scale vertical stratification and guild composition of saproxylic beetles in lowland and montane forests: Similar patterns despite low faunal overlap. PLoS ONE . 11: e0149506.
Wermelinger, B., Flückiger, P. F., Obrist, M. K., and Duelli, P. 2007. Horizontal and vertical distribution of saproxylic beetles (Col., Buprestidae, Cerambycidae, Scolytinae) across sections of forest edges.J. Appl .Entomol. 131 : 104-114.
Yusah, K. M., and Foster, W. A. 2016. Tree size and habitat complexity affect ant communities (Hymenoptera: Formicidae) in the high canopy of Bornean rain forest. Myrmecol. News 23 : 15-23.
Yusah, K. M., Foster, W. A., Reynolds, G., and Fayle, T. M. 2018. Ant mosaics in Bornean primary rain forest high canopy depend on spatial scale, time of day, and sampling method. PeerJ . 6 : e4231.
Zellweger, F., Roth, T., Bugmann, H., and Bollmann, K. 2017. Beta diversity of plants, birds and butterflies is closely associated with climate and habitat structure. Global Ecol. Biogeogr.26 : 898-906.