AUTHOR CONTRIBUTIONS

N.L., S.B. and N.D. designed the experiment. F.S., N.L., A.V. and N.D. organized sampling expeditions. F.S. and N.L. sampled fish during field expeditions. N.L. processed samples in the laboratory (fish dissections and DNA extractions). N.L. and S.B. performed bioinformatic analyses. N.L. wrote the manuscript. All authors reviewed the manuscript.