Author Contributions
Sarah C. Deckel contributed to conceptual design of the study, organizing, and executing the data collection/field season, all analysis (including interpretation), and writing the draft for this manuscript. William V. DeLuca assisted with conceptualization of the study design, developing field methods and field work, and contributed to final review and editing of the manuscript. Alexander R. Gerson contributed expertise to the manuscript draft and analysis interpretation, as well conceptualization of the study design. David I. King contributed to input on data collection methods, funding opportunities, conceptualization of study design, as well as expertise on the manuscript drafts. All authors gave final approval for this version to be published.