Fig 1 . Study locations coded by biomes. FGS (Flooded Grasslands and Savannas); TSCF (Tropical and Sub-tropical Coniferous Forests); TSDBF (Tropical and Sub-tropical Dry Broadleaf Forests); TSGSS (Tropical and Sub-tropical Grasslands, Shrublands and Savannas); and TSMBF (Tropical and Sub-tropical Moist Broadleaf Forests). The dotted lines show the boundary of the sub-tropical zone